*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }

:root {
  --color-bg: #f5f0e8;
  --color-bg-alt: #ede8dc;
  --color-dark: #1a2332;
  --color-dark-alt: #2c3e50;
  --color-accent: #2e7d52;
  --color-accent-light: #4caf78;
  --color-accent-warm: #c0722a;
  --color-text: #333d45;
  --color-text-light: #5a6472;
  --color-border: #d4cdbf;
  --color-white: #ffffff;
  --font-display: 'Playfair Display', serif;
  --font-body: 'Lato', sans-serif;
  --font-serif: 'Merriweather', serif;
  --font-ui: 'Raleway', sans-serif;
  --radius: 8px;
  --radius-lg: 16px;
  --shadow: 0 4px 20px rgba(0,0,0,0.08);
  --shadow-lg: 0 8px 40px rgba(0,0,0,0.14);
  --transition: 0.3s ease;
}

html { scroll-behavior: smooth; font-size: 16px; }
body { font-family: var(--font-body); background: var(--color-bg); color: var(--color-text); line-height: 1.7; }
img { max-width: 100%; height: auto; display: block; }
a { color: var(--color-accent); text-decoration: none; transition: color var(--transition); }
a:hover { color: var(--color-accent-light); }
ul { list-style: none; }

#page-loader {
  position: fixed; inset: 0; background: var(--color-dark);
  display: flex; align-items: center; justify-content: center;
  z-index: 9999; transition: opacity 0.4s ease;
}
#page-loader.hidden { opacity: 0; pointer-events: none; }
.loader-inner { text-align: center; color: var(--color-white); }
.loader-feather {
  width: 48px; height: 48px; margin: 0 auto 16px;
  border: 3px solid rgba(255,255,255,0.2);
  border-top-color: var(--color-accent-light);
  border-radius: 50%;
  animation: spin 0.9s linear infinite;
}
@keyframes spin { to { transform: rotate(360deg); } }
.loader-inner span { font-family: var(--font-ui); font-size: 0.85rem; letter-spacing: 0.1em; opacity: 0.7; }

.site-header {
  position: sticky; top: 0; z-index: 100;
  background: rgba(26, 35, 50, 0.97);
  backdrop-filter: blur(8px);
  border-bottom: 1px solid rgba(255,255,255,0.06);
  transition: box-shadow var(--transition);
}
.site-header.scrolled { box-shadow: 0 2px 20px rgba(0,0,0,0.3); }
.header-inner {
  max-width: 1200px; margin: 0 auto;
  padding: 0 24px;
  display: flex; align-items: center; justify-content: space-between;
  height: 68px;
}
.site-logo { display: flex; align-items: center; gap: 12px; text-decoration: none; }
.site-logo img { border-radius: 50%; border: 2px solid var(--color-accent); }
.logo-text { display: flex; flex-direction: column; }
.logo-title { font-family: var(--font-display); font-size: 1.1rem; font-weight: 700; color: var(--color-white); line-height: 1.2; }
.logo-sub { font-family: var(--font-ui); font-size: 0.65rem; color: var(--color-accent-light); letter-spacing: 0.08em; text-transform: uppercase; }

.main-nav ul { display: flex; align-items: center; gap: 6px; }
.main-nav a {
  font-family: var(--font-ui); font-size: 0.88rem; font-weight: 500;
  color: rgba(255,255,255,0.82); padding: 8px 14px;
  border-radius: var(--radius); transition: all var(--transition);
  letter-spacing: 0.03em;
}
.main-nav a:hover, .main-nav a.active {
  color: var(--color-white); background: rgba(255,255,255,0.1);
}
.has-dropdown { position: relative; }
.dropdown-menu {
  position: absolute; top: calc(100% + 8px); left: 50%; transform: translateX(-50%);
  background: var(--color-dark); border: 1px solid rgba(255,255,255,0.1);
  border-radius: var(--radius); min-width: 220px;
  box-shadow: var(--shadow-lg); opacity: 0; visibility: hidden;
  transition: all var(--transition); pointer-events: none;
  padding: 8px 0;
}
.has-dropdown:hover .dropdown-menu { opacity: 1; visibility: visible; pointer-events: all; }
.dropdown-menu li a {
  display: block; padding: 10px 20px; border-radius: 0;
  font-size: 0.85rem; color: rgba(255,255,255,0.8);
}
.dropdown-menu li a:hover { background: rgba(255,255,255,0.08); color: var(--color-white); }
.arrow { font-size: 0.6rem; margin-left: 4px; vertical-align: middle; }

.burger-btn {
  display: none; flex-direction: column; justify-content: center;
  gap: 5px; background: none; border: none; cursor: pointer; padding: 8px;
}
.burger-btn span { display: block; width: 24px; height: 2px; background: var(--color-white); border-radius: 2px; transition: all var(--transition); }

.mobile-menu {
  position: fixed; top: 0; right: -100%; width: min(320px, 90vw); height: 100vh;
  background: var(--color-dark-alt); z-index: 200;
  transition: right 0.35s cubic-bezier(0.4,0,0.2,1);
  padding: 72px 24px 32px; overflow-y: auto;
}
.mobile-menu.open { right: 0; }
.mobile-menu-close {
  position: absolute; top: 18px; right: 18px;
  background: none; border: none; color: var(--color-white);
  font-size: 1.8rem; cursor: pointer; line-height: 1; padding: 4px;
}
.mobile-menu ul { display: flex; flex-direction: column; gap: 4px; }
.mobile-menu a { color: rgba(255,255,255,0.85); font-family: var(--font-ui); font-size: 1rem; padding: 12px 8px; display: block; border-bottom: 1px solid rgba(255,255,255,0.07); }
.mobile-menu a:hover { color: var(--color-accent-light); }
.mobile-section-title { color: var(--color-accent-light); font-family: var(--font-ui); font-size: 0.75rem; text-transform: uppercase; letter-spacing: 0.12em; padding: 16px 8px 4px; }
.mobile-overlay {
  position: fixed; inset: 0; background: rgba(0,0,0,0.5);
  z-index: 199; opacity: 0; visibility: hidden; transition: all var(--transition);
}
.mobile-overlay.visible { opacity: 1; visibility: visible; }

.hero-section {
  min-height: 90vh; display: grid; grid-template-columns: 1fr 1fr;
  align-items: center; max-width: 1200px; margin: 0 auto;
  padding: 80px 24px 60px; gap: 60px;
}
.hero-content { order: 1; }
.hero-eyebrow {
  font-family: var(--font-ui); font-size: 0.78rem; font-weight: 600;
  color: var(--color-accent); text-transform: uppercase; letter-spacing: 0.15em;
  display: flex; align-items: center; gap: 10px; margin-bottom: 20px;
}
.hero-eyebrow::before { content: ''; display: block; width: 32px; height: 2px; background: var(--color-accent); }
.hero-title {
  font-family: var(--font-display); font-size: clamp(2.4rem, 5vw, 3.8rem);
  font-weight: 700; line-height: 1.15; color: var(--color-dark); margin-bottom: 24px;
}
.hero-title em { font-style: italic; color: var(--color-accent); }
.hero-desc { font-family: var(--font-serif); font-size: 1.05rem; color: var(--color-text-light); margin-bottom: 36px; line-height: 1.85; }
.hero-cta { display: flex; gap: 16px; flex-wrap: wrap; }
.btn-primary {
  display: inline-flex; align-items: center; gap: 8px;
  background: var(--color-accent); color: var(--color-white);
  padding: 14px 28px; border-radius: var(--radius);
  font-family: var(--font-ui); font-weight: 600; font-size: 0.9rem;
  letter-spacing: 0.04em; transition: all var(--transition);
  border: 2px solid var(--color-accent);
}
.btn-primary:hover { background: var(--color-accent-light); border-color: var(--color-accent-light); color: var(--color-white); transform: translateY(-2px); box-shadow: 0 6px 20px rgba(46,125,82,0.3); }
.btn-secondary {
  display: inline-flex; align-items: center; gap: 8px;
  background: transparent; color: var(--color-dark);
  padding: 14px 28px; border-radius: var(--radius);
  font-family: var(--font-ui); font-weight: 600; font-size: 0.9rem;
  letter-spacing: 0.04em; transition: all var(--transition);
  border: 2px solid var(--color-dark);
}
.btn-secondary:hover { background: var(--color-dark); color: var(--color-white); transform: translateY(-2px); }

.hero-image { order: 2; position: relative; }
.hero-image-wrap {
  position: relative; border-radius: var(--radius-lg);
  overflow: hidden; box-shadow: var(--shadow-lg);
  aspect-ratio: 4/5;
}
.hero-image-wrap img { width: 100%; height: 100%; object-fit: cover; }
.hero-image-badge {
  position: absolute; bottom: 24px; left: 24px;
  background: rgba(26,35,50,0.88); backdrop-filter: blur(4px);
  color: var(--color-white); padding: 12px 18px;
  border-radius: var(--radius); border-left: 3px solid var(--color-accent-light);
}
.hero-image-badge strong { display: block; font-family: var(--font-display); font-size: 1rem; }
.hero-image-badge span { font-size: 0.78rem; opacity: 0.75; font-family: var(--font-ui); }

.section-header { text-align: center; margin-bottom: 56px; }
.section-label {
  font-family: var(--font-ui); font-size: 0.75rem; font-weight: 600;
  color: var(--color-accent); text-transform: uppercase; letter-spacing: 0.15em;
  display: block; margin-bottom: 12px;
}
.section-title { font-family: var(--font-display); font-size: clamp(1.8rem, 3.5vw, 2.8rem); font-weight: 700; color: var(--color-dark); line-height: 1.2; }
.section-desc { font-family: var(--font-serif); font-size: 1rem; color: var(--color-text-light); max-width: 600px; margin: 16px auto 0; line-height: 1.8; font-style: italic; }

.featured-section { padding: 80px 24px; background: var(--color-bg-alt); }
.featured-inner { max-width: 1200px; margin: 0 auto; }
.featured-grid { display: grid; grid-template-columns: 1fr 1fr; gap: 32px; }
.featured-card {
  background: var(--color-white); border-radius: var(--radius-lg);
  overflow: hidden; box-shadow: var(--shadow);
  transition: transform var(--transition), box-shadow var(--transition);
  display: flex; flex-direction: column;
}
.featured-card:hover { transform: translateY(-6px); box-shadow: var(--shadow-lg); }
.featured-card.large { grid-column: span 2; display: grid; grid-template-columns: 55% 45%; }
.card-img { overflow: hidden; }
.card-img img { width: 100%; height: 100%; object-fit: cover; transition: transform 0.5s ease; }
.featured-card:hover .card-img img { transform: scale(1.04); }
.featured-card:not(.large) .card-img { height: 240px; }
.card-body { padding: 28px 32px; display: flex; flex-direction: column; justify-content: center; }
.card-tag {
  display: inline-block; font-family: var(--font-ui); font-size: 0.72rem;
  font-weight: 700; text-transform: uppercase; letter-spacing: 0.1em;
  color: var(--color-accent); background: rgba(46,125,82,0.1);
  padding: 4px 12px; border-radius: 20px; margin-bottom: 14px;
}
.card-title { font-family: var(--font-display); font-size: 1.45rem; font-weight: 700; color: var(--color-dark); margin-bottom: 12px; line-height: 1.3; }
.card-title a { color: inherit; }
.card-title a:hover { color: var(--color-accent); }
.card-excerpt { font-family: var(--font-serif); font-size: 0.9rem; color: var(--color-text-light); line-height: 1.8; margin-bottom: 20px; font-style: italic; }
.card-meta { display: flex; align-items: center; gap: 16px; font-family: var(--font-ui); font-size: 0.78rem; color: var(--color-text-light); margin-top: auto; padding-top: 16px; border-top: 1px solid var(--color-border); }
.card-link { font-family: var(--font-ui); font-size: 0.85rem; font-weight: 600; color: var(--color-accent); display: inline-flex; align-items: center; gap: 6px; margin-top: 16px; }
.card-link:hover { color: var(--color-accent-light); gap: 10px; }

.info-section { padding: 80px 24px; max-width: 1200px; margin: 0 auto; }
.info-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 32px; }
.info-card {
  background: var(--color-white); border-radius: var(--radius-lg);
  padding: 36px 28px; text-align: center;
  border-top: 4px solid var(--color-accent);
  box-shadow: var(--shadow); transition: transform var(--transition);
}
.info-card:hover { transform: translateY(-4px); }
.info-icon { font-size: 2.4rem; margin-bottom: 16px; }
.info-card h3 { font-family: var(--font-display); font-size: 1.2rem; font-weight: 700; color: var(--color-dark); margin-bottom: 12px; }
.info-card p { font-family: var(--font-serif); font-size: 0.88rem; color: var(--color-text-light); line-height: 1.8; font-style: italic; }

.species-section { padding: 80px 24px; background: var(--color-dark); }
.species-inner { max-width: 1200px; margin: 0 auto; }
.species-section .section-title { color: var(--color-white); }
.species-section .section-desc { color: rgba(255,255,255,0.6); }
.species-section .section-label { color: var(--color-accent-light); }
.species-grid { display: grid; grid-template-columns: repeat(4, 1fr); gap: 20px; }
.species-card {
  position: relative; border-radius: var(--radius); overflow: hidden;
  aspect-ratio: 3/4; cursor: pointer;
  box-shadow: var(--shadow);
}
.species-card img { width: 100%; height: 100%; object-fit: cover; transition: transform 0.5s ease; }
.species-card:hover img { transform: scale(1.06); }
.species-overlay {
  position: absolute; inset: 0;
  background: linear-gradient(to top, rgba(26,35,50,0.9) 40%, transparent 70%);
  display: flex; flex-direction: column; justify-content: flex-end;
  padding: 20px; transition: background var(--transition);
}
.species-card:hover .species-overlay { background: linear-gradient(to top, rgba(26,35,50,0.95) 50%, rgba(26,35,50,0.2) 80%); }
.species-name { font-family: var(--font-display); font-size: 1rem; font-weight: 700; color: var(--color-white); margin-bottom: 4px; }
.species-latin { font-family: var(--font-serif); font-size: 0.75rem; color: rgba(255,255,255,0.6); font-style: italic; margin-bottom: 8px; }
.species-link { font-family: var(--font-ui); font-size: 0.75rem; color: var(--color-accent-light); font-weight: 600; text-transform: uppercase; letter-spacing: 0.08em; opacity: 0; transition: opacity var(--transition); }
.species-card:hover .species-link { opacity: 1; }

.page-hero {
  background: var(--color-dark); padding: 80px 24px 60px;
  text-align: center; position: relative; overflow: hidden;
}
.page-hero::before {
  content: ''; position: absolute; inset: 0;
  background: linear-gradient(135deg, rgba(46,125,82,0.15) 0%, transparent 60%);
}
.page-hero-inner { max-width: 800px; margin: 0 auto; position: relative; }
.page-hero .breadcrumb {
  font-family: var(--font-ui); font-size: 0.78rem; color: rgba(255,255,255,0.5);
  margin-bottom: 20px; display: flex; justify-content: center; gap: 8px; align-items: center;
}
.page-hero .breadcrumb a { color: rgba(255,255,255,0.6); }
.page-hero .breadcrumb a:hover { color: var(--color-accent-light); }
.page-hero .breadcrumb span { color: rgba(255,255,255,0.3); }
.page-hero h1 { font-family: var(--font-display); font-size: clamp(2rem, 5vw, 3.2rem); font-weight: 700; color: var(--color-white); line-height: 1.2; margin-bottom: 20px; }
.page-hero .hero-meta {
  display: flex; align-items: center; justify-content: center; gap: 20px;
  font-family: var(--font-ui); font-size: 0.8rem; color: rgba(255,255,255,0.55);
}
.page-hero .hero-meta span { display: flex; align-items: center; gap: 6px; }

.article-layout { max-width: 1200px; margin: 0 auto; padding: 60px 24px; display: grid; grid-template-columns: 1fr 340px; gap: 60px; align-items: start; }
.article-content { min-width: 0; }
.article-lead-image { border-radius: var(--radius-lg); overflow: hidden; margin-bottom: 48px; box-shadow: var(--shadow-lg); }
.article-lead-image img { width: 100%; height: 480px; object-fit: cover; }
.article-lead-image figcaption { background: var(--color-bg-alt); padding: 12px 20px; font-family: var(--font-ui); font-size: 0.78rem; color: var(--color-text-light); text-align: center; }
.article-body h2 { font-family: var(--font-display); font-size: 1.7rem; font-weight: 700; color: var(--color-dark); margin: 40px 0 18px; line-height: 1.3; }
.article-body h3 { font-family: var(--font-display); font-size: 1.25rem; font-weight: 600; color: var(--color-dark-alt); margin: 28px 0 12px; }
.article-body p { font-family: var(--font-serif); font-size: 1rem; color: var(--color-text); line-height: 1.9; margin-bottom: 20px; }
.article-body p:first-child { font-size: 1.1rem; color: var(--color-text-light); font-style: italic; }
.article-body ul, .article-body ol { margin: 0 0 20px 24px; list-style: disc; }
.article-body li { font-family: var(--font-serif); font-size: 0.95rem; color: var(--color-text); line-height: 1.8; margin-bottom: 8px; }
.article-body a { color: var(--color-accent); text-decoration: underline; text-underline-offset: 3px; }
.article-body a:hover { color: var(--color-accent-light); }
.article-inline-img { border-radius: var(--radius); overflow: hidden; margin: 32px 0; box-shadow: var(--shadow); }
.article-inline-img img { width: 100%; height: 320px; object-fit: cover; }
.article-inline-img figcaption { background: var(--color-bg-alt); padding: 10px 16px; font-family: var(--font-ui); font-size: 0.75rem; color: var(--color-text-light); }
.article-callout {
  background: var(--color-bg-alt); border-left: 4px solid var(--color-accent);
  border-radius: 0 var(--radius) var(--radius) 0;
  padding: 20px 24px; margin: 28px 0;
}
.article-callout p { font-family: var(--font-serif); font-style: italic; color: var(--color-text-light); margin: 0; font-size: 0.95rem; }
.article-sources { margin-top: 48px; padding-top: 32px; border-top: 2px solid var(--color-border); }
.article-sources h4 { font-family: var(--font-ui); font-size: 0.85rem; text-transform: uppercase; letter-spacing: 0.1em; color: var(--color-text-light); margin-bottom: 12px; }
.article-sources ul { list-style: none; margin: 0; }
.article-sources li { font-family: var(--font-ui); font-size: 0.82rem; color: var(--color-text-light); margin-bottom: 8px; padding-left: 16px; position: relative; }
.article-sources li::before { content: '—'; position: absolute; left: 0; color: var(--color-accent); }
.article-updated { font-family: var(--font-ui); font-size: 0.78rem; color: var(--color-text-light); margin-top: 32px; padding-top: 16px; border-top: 1px solid var(--color-border); }

.sidebar { position: sticky; top: 88px; }
.sidebar-widget { background: var(--color-white); border-radius: var(--radius-lg); padding: 28px; margin-bottom: 24px; box-shadow: var(--shadow); }
.sidebar-widget h3 { font-family: var(--font-display); font-size: 1.05rem; font-weight: 700; color: var(--color-dark); margin-bottom: 16px; padding-bottom: 12px; border-bottom: 2px solid var(--color-border); }
.sidebar-nav ul { display: flex; flex-direction: column; gap: 4px; }
.sidebar-nav li a { display: block; padding: 8px 12px; border-radius: var(--radius); font-family: var(--font-ui); font-size: 0.85rem; color: var(--color-text); transition: all var(--transition); border-left: 3px solid transparent; }
.sidebar-nav li a:hover, .sidebar-nav li a.active { color: var(--color-accent); background: rgba(46,125,82,0.07); border-left-color: var(--color-accent); }
.sidebar-fact { font-family: var(--font-serif); font-size: 0.88rem; color: var(--color-text-light); line-height: 1.7; font-style: italic; }
.sidebar-fact strong { font-style: normal; color: var(--color-dark); font-family: var(--font-ui); display: block; margin-bottom: 6px; }

.related-section { background: var(--color-bg-alt); padding: 60px 24px; }
.related-inner { max-width: 1200px; margin: 0 auto; }
.related-grid { display: grid; grid-template-columns: repeat(3, 1fr); gap: 24px; }
.related-card {
  background: var(--color-white); border-radius: var(--radius-lg); overflow: hidden;
  box-shadow: var(--shadow); transition: transform var(--transition), box-shadow var(--transition);
}
.related-card:hover { transform: translateY(-4px); box-shadow: var(--shadow-lg); }
.related-card-img { height: 180px; overflow: hidden; }
.related-card-img img { width: 100%; height: 100%; object-fit: cover; transition: transform 0.5s ease; }
.related-card:hover .related-card-img img { transform: scale(1.05); }
.related-card-body { padding: 20px; }
.related-card-tag { font-family: var(--font-ui); font-size: 0.7rem; font-weight: 700; text-transform: uppercase; letter-spacing: 0.1em; color: var(--color-accent); margin-bottom: 8px; display: block; }
.related-card-title { font-family: var(--font-display); font-size: 1.05rem; font-weight: 700; color: var(--color-dark); line-height: 1.3; margin-bottom: 8px; }
.related-card-title a { color: inherit; }
.related-card-title a:hover { color: var(--color-accent); }
.related-card-desc { font-family: var(--font-serif); font-size: 0.82rem; color: var(--color-text-light); line-height: 1.7; font-style: italic; }

.simple-page { max-width: 860px; margin: 0 auto; padding: 60px 24px; }
.simple-page h2 { font-family: var(--font-display); font-size: 1.6rem; font-weight: 700; color: var(--color-dark); margin: 36px 0 16px; }
.simple-page h3 { font-family: var(--font-display); font-size: 1.2rem; font-weight: 600; color: var(--color-dark-alt); margin: 24px 0 10px; }
.simple-page p, .simple-page li { font-family: var(--font-serif); font-size: 0.95rem; color: var(--color-text); line-height: 1.9; margin-bottom: 16px; }
.simple-page ul { margin-left: 24px; list-style: disc; }
.simple-page a { color: var(--color-accent); text-decoration: underline; }

.contact-layout { max-width: 900px; margin: 0 auto; padding: 60px 24px; }
.contact-grid { display: grid; grid-template-columns: 1fr 1fr; gap: 48px; }
.contact-info h2 { font-family: var(--font-display); font-size: 1.5rem; font-weight: 700; color: var(--color-dark); margin-bottom: 16px; }
.contact-info p { font-family: var(--font-serif); font-size: 0.92rem; color: var(--color-text-light); line-height: 1.8; margin-bottom: 16px; font-style: italic; }
.contact-email-block { background: var(--color-bg-alt); border-radius: var(--radius); padding: 20px 24px; border-left: 4px solid var(--color-accent); margin-top: 24px; }
.contact-email-block h4 { font-family: var(--font-ui); font-size: 0.8rem; text-transform: uppercase; letter-spacing: 0.1em; color: var(--color-text-light); margin-bottom: 8px; }
.contact-email-link { font-family: var(--font-display); font-size: 1.1rem; font-weight: 600; color: var(--color-accent); }
.contact-email-link:hover { color: var(--color-accent-light); }
.contact-details h3 { font-family: var(--font-display); font-size: 1.2rem; font-weight: 700; color: var(--color-dark); margin-bottom: 20px; }
.contact-detail-item { display: flex; gap: 16px; margin-bottom: 20px; align-items: flex-start; }
.detail-icon { width: 40px; height: 40px; background: rgba(46,125,82,0.1); border-radius: 50%; display: flex; align-items: center; justify-content: center; font-size: 1.1rem; flex-shrink: 0; }
.detail-text strong { display: block; font-family: var(--font-ui); font-size: 0.82rem; text-transform: uppercase; letter-spacing: 0.08em; color: var(--color-text-light); margin-bottom: 4px; }
.detail-text span, .detail-text a { font-family: var(--font-body); font-size: 0.92rem; color: var(--color-text); }

.site-footer {
  background: var(--color-dark); color: rgba(255,255,255,0.75);
  padding: 64px 24px 32px;
}
.footer-inner { max-width: 1200px; margin: 0 auto; }
.footer-grid { display: grid; grid-template-columns: 2fr 1fr 1fr 1.5fr; gap: 48px; margin-bottom: 48px; }
.footer-brand { }
.footer-logo { display: flex; align-items: center; gap: 12px; text-decoration: none; margin-bottom: 16px; }
.footer-logo img { border-radius: 50%; border: 2px solid var(--color-accent); }
.footer-logo span { font-family: var(--font-display); font-size: 1rem; font-weight: 700; color: var(--color-white); }
.footer-brand p { font-family: var(--font-serif); font-size: 0.85rem; color: rgba(255,255,255,0.5); line-height: 1.8; font-style: italic; }
.footer-links h4 { font-family: var(--font-ui); font-size: 0.78rem; text-transform: uppercase; letter-spacing: 0.12em; color: rgba(255,255,255,0.4); margin-bottom: 16px; }
.footer-links ul { display: flex; flex-direction: column; gap: 8px; }
.footer-links a { font-family: var(--font-body); font-size: 0.88rem; color: rgba(255,255,255,0.65); transition: color var(--transition); }
.footer-links a:hover { color: var(--color-accent-light); }
.footer-contact h4 { font-family: var(--font-ui); font-size: 0.78rem; text-transform: uppercase; letter-spacing: 0.12em; color: rgba(255,255,255,0.4); margin-bottom: 16px; }
.footer-contact p { font-family: var(--font-serif); font-size: 0.83rem; color: rgba(255,255,255,0.5); margin-bottom: 10px; font-style: italic; }
.footer-email { font-family: var(--font-ui); font-size: 0.9rem; color: var(--color-accent-light); font-weight: 600; }
.footer-email:hover { color: var(--color-white); }
.footer-sources { margin-top: 20px; padding-top: 16px; border-top: 1px solid rgba(255,255,255,0.08); }
.footer-sources h5 { font-family: var(--font-ui); font-size: 0.72rem; text-transform: uppercase; letter-spacing: 0.1em; color: rgba(255,255,255,0.35); margin-bottom: 10px; }
.footer-sources ul { display: flex; flex-direction: column; gap: 6px; }
.footer-sources a { font-family: var(--font-body); font-size: 0.78rem; color: rgba(255,255,255,0.45); }
.footer-sources a:hover { color: var(--color-accent-light); }
.footer-bottom { border-top: 1px solid rgba(255,255,255,0.08); padding-top: 24px; display: flex; justify-content: space-between; align-items: center; flex-wrap: wrap; gap: 12px; }
.footer-bottom p { font-family: var(--font-ui); font-size: 0.78rem; color: rgba(255,255,255,0.35); }
.footer-bottom a { color: rgba(255,255,255,0.45); }
.footer-bottom a:hover { color: var(--color-accent-light); }

.cookie-banner {
  position: fixed; bottom: -100px; left: 50%; transform: translateX(-50%);
  width: min(680px, calc(100vw - 32px));
  background: var(--color-dark); border: 1px solid rgba(255,255,255,0.1);
  border-radius: var(--radius-lg); box-shadow: var(--shadow-lg);
  padding: 20px 24px; z-index: 500;
  transition: bottom 0.4s cubic-bezier(0.4,0,0.2,1);
  display: flex; align-items: center;
}
.cookie-banner.visible { bottom: 24px; }
.cookie-content { display: flex; align-items: center; gap: 20px; width: 100%; flex-wrap: wrap; }
.cookie-content p { font-family: var(--font-body); font-size: 0.85rem; color: rgba(255,255,255,0.75); line-height: 1.6; flex: 1; min-width: 200px; }
.cookie-content a { color: var(--color-accent-light); text-decoration: underline; }
.cookie-buttons { display: flex; gap: 10px; flex-shrink: 0; }
.btn-cookie {
  padding: 9px 22px; border-radius: var(--radius); font-family: var(--font-ui);
  font-size: 0.82rem; font-weight: 600; cursor: pointer; transition: all var(--transition);
  border: 2px solid transparent; letter-spacing: 0.04em;
}
.btn-accept { background: var(--color-accent); color: var(--color-white); border-color: var(--color-accent); }
.btn-accept:hover { background: var(--color-accent-light); border-color: var(--color-accent-light); }
.btn-reject { background: transparent; color: rgba(255,255,255,0.7); border-color: rgba(255,255,255,0.2); }
.btn-reject:hover { border-color: rgba(255,255,255,0.5); color: var(--color-white); }

@media (max-width: 1024px) {
  .featured-grid { grid-template-columns: 1fr; }
  .featured-card.large { grid-column: span 1; grid-template-columns: 1fr; }
  .featured-card.large .card-img { height: 300px; }
  .footer-grid { grid-template-columns: 1fr 1fr; gap: 32px; }
  .species-grid { grid-template-columns: repeat(2, 1fr); }
  .article-layout { grid-template-columns: 1fr; }
  .sidebar { position: static; }
  .hero-section { grid-template-columns: 1fr; min-height: auto; }
  .hero-image { order: -1; }
  .hero-image-wrap { aspect-ratio: 16/9; }
  .info-grid { grid-template-columns: 1fr 1fr; }
}

@media (max-width: 768px) {
  .main-nav { display: none; }
  .burger-btn { display: flex; }
  .hero-title { font-size: 2.2rem; }
  .related-grid { grid-template-columns: 1fr 1fr; }
  .contact-grid { grid-template-columns: 1fr; }
  .footer-grid { grid-template-columns: 1fr; gap: 28px; }
  .info-grid { grid-template-columns: 1fr; }
  .species-grid { grid-template-columns: repeat(2, 1fr); }
}

@media (max-width: 500px) {
  .related-grid { grid-template-columns: 1fr; }
  .hero-cta { flex-direction: column; }
  .btn-primary, .btn-secondary { text-align: center; justify-content: center; }
  .species-grid { grid-template-columns: 1fr 1fr; }
  .cookie-content { flex-direction: column; align-items: flex-start; }
}